为此，我替换了：
if project.pk:
project.userprofile_set = self.cleaned_data['userprofiles']
self.save_m2m()
通过：
if project.pk:
# Get the existing relationships
current_project_selections = Membership.objects.filter(project=project)
current_selections = [o.userprofile for o in current_project_selections]
# Get the submitted relationships
submitted_selections = self.cleaned_data['userprofiles']
# Create new relation in Membership table if they do not exist
for userprofile in submitted_selections :
if userprofile not in current_selections:
Membership(project=project,userprofile=userprofile).save()
# Remove the relations that were deselected from the Membership table
for project_userprofile in current_project_selections:
if project_userprofile.userprofile not in submitted_selections :
project_userprofile.delete()
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